Alemseged Lab Publications
Read about our research in any of the publications listed below.Full Publication List
2025
Lawrence, A., Hammond, A., Ward, C.V. Acetabular orientation, pelvic shape, and the evolution of hominin bipedality. J. Hum. Evol. 200, 103633. Online Article
2024
Negash, E.W., Alemseged, Z., Barr, W.A., Behrensmeyer, A.K., Blumenthal, S.A., Bobe, R., Carvalho, S., Cerling, T.E., Chritz, K.L., McGuire, E., Uno, K.T., Wood, B., Wynn, J.G. Modern African ecosystems as landscape-scale analogues for reconstructing woody cover and early hominin environments. J. Hum. Evol. 197, 103604. Online Article
Alene, M. & Alemseged, Z. Volcanism, Tectonics and Hominin Evolution in the Afar Rift, East Africa. SGAT Bull. 25(2): 105-107.
Davies, T.W., Gunz, P., Spoor, F., Alemseged, Z., Gidna, A., Hublin, J.-J., Kimbel, W.H., Kullmer, O., Plummer, W.P., Zanolli, C., Skinner, M.M., 2024. Dental morphology in Homo habilis and its implications for the evolution of early Homo. Nat Commun. 15, 286. Online Article
2023
Alemseged, Z., 2023. Yves Coppens: Scientist, educator, popularizer, and thinker. Bull. Mus. Anthropol. préhist. Monaco, 10: 47-51.
Bobe, R., Aldeias, V., Alemseged, Z., Anemone, R.L., Archer, W., Aumaître, G., Bamford, M.K., Biro, D., Bourlès, D.L., Doyle Boyd, M., Braun, D.R., Capelli, C., d’Oliveira Coelho, J., Habermann, J.M., Head, J.J., Keddadouche, K., Kupczik, K., Lebatard, A.-E., Lüdecke, T., Macôa, A., Martínez, F.I., Mathe, J., Mendes, C., Paulo, L.M., Pinto, M., Presnyakova, D., Püschel, T.A., Regala, F.T., Sier, M., Ferreira Da Silva, M.J., Stalmans, M., Carvalho, S., 2023. The first Miocene fossils from coastal woodlands in the southern East African Rift. iScience. 26, 107644. Online Article
Alemseged, Z., 2023. Reappraising the palaeobiology of Australopithecus. Nature. 617, 45–54. Online Article
Stamos, P.A., Alemseged, Z., 2023. Hominin locomotion and evolution in the Late Miocene to Late Pliocene. J. Hum. Evol. 178, 103332. Online Article
Smith, A.L., Davis, J., Panagiotopoulou, O., Taylor, A.B., Robinson, C., Ward, C.V., Kimbel, W.H., Alemseged, Z., Ross, C.F., 2023. Does the model reflect the system? When two-dimensional biomechanics is not ‘good enough.’ J. R. Soc. Interface. 20, 20220536. Online Article
2022
Geraads, D. & Alemseged, Z. Comment (Case 3847) – Opposition to the proposed conservation of Simopithecus oswaldi Andrews, 1916 (currently Theropithecus oswaldi; Mammalia, Primates, Cercopithecidae), by reversal of precedence with Cynocephalus atlanticus Thomas, 1884 (see BZN 78: 99–106 [Case]). The Bulletin of Zoological Nomenclature 79. Online Article
Negash, E.W., Bobe, R., Alemseged, Z., Wynn, J. Mammalian Diversity Patterns and Paleoecology in the Lower Omo Valley, Ethiopia. In: Reynolds, S.C., Bobe, R. (Eds.), African Paleoecology and Human Evolution. Cambridge University Press, pp. 289–297. Online Article
Bobe, R., Geraads, D., Wynn, J.G., Reed, D., Barr, A.W., Alemseged, Z. Fossil Vertebrates and Paleoenvironments of the Pliocene Hadar Formation at Dikika, Ethiopia. In: Reynolds, S.C., Bobe, R. (Eds.), African Paleoecology and Human Evolution. Cambridge University Press, pp. 229–241. Online Article
2021
Alemseged, Z. Earliest Stone Tool Use in Hominins and the Significance of the Dikika Cutmarks: Beyond the Cuts! In: Coppens, Y., Vialet, A. (Eds.), Un Bouquet d’ancêtres. Premiers Humains: Qui Était Qui, Qui a Fait Quoi, Où et Quand? Nouvelle Impremerie Laballery, Clamecy, France, pp. 101–122.
Smith, A.L., Robinson, C., Taylor, A.B., Panagiotopoulou, O., Davis, J., Ward, C., Kimbel, W.H., Alemseged, Z., Ross, C.F. Comparative biomechanics of the Pan and Macaca mandibles during mastication: finite element modelling of loading, deformation and strain regimes. Interface Focus 11:20210031. Online Article
Geraads, D., Reed, D., Barr, W.A., Bobe, R., Stamos, P., Alemseged, Z. Plio-Pleistocene mammals from Mille-Logya, Ethiopia, and the post-Hadar faunal change. J. Quat. Sci. 2021:1-17. Online article
Du, A., Rowan, J., Wang, S.C., Wood, B.A., Alemseged, Z., On fossil recovery potential in the Australopithecus anamensis-Australopithecus afarensis lineage: A reply to Žliobaitė (2020). Journal of Human Evolution. 157(2021). Online Article
Davies, T.W., Alemseged, Z., Gidna, A., Jean-Jacques, H., Kimbel, W.H., Kullmer, O., Spoor, F., Zanolli, C., Skinner, M.M. Accessory cusp expression at the enamel-dentine junction of hominin mandibular molars. PeerJ 9:e11415 Online Article
2020
Wynn, J.G., Alemseged, Z., Bobe, R., Grine, F.E., Negash, E.W., Sponheimer, M. Isotopic evidence for the timing of the dietary shift toward C4 foods in eastern African Paranthropus. PNAS. DOI: 10.1073/pnas.2006221117 Online Article
Negash, E.W., Alemseged, Z., Bobe, R., Grine, F.E., Sponheimer, M., Wynn, J.G. Dietary trends in herbivores from the Shungura Formation, southwestern Ethiopia. PNAS DOI: 10.1073/pnas.2006982117 Online Article
Alemseged, Z., Wynn, J.G., Geraads, D., Reed, D., Barr, W.A., Bobe, R., McPherron, S.P., Deino, A., Alene, M., Sier, M.J., Roman, D., Mohan, J. Fossils from Mille-Logya, Afar, Ethiopia, elucidate the link between Pliocene environmental changes and Homo origins. Nature Communications. (2020)11:2480. Online Article
Gunz, P., Neubauer, S., Faulk, D., Tafforeau, P., Le Cabec, A., Smith, T.M., Kimbel, W.H., Spoor, F., Alemseged, Z., Australopithecus afarensis endocasts suggest ape-like brain organization and prolonged brain growth. Science Advances. 6: eaaz4729. Online Article
Lockey, A., Alemseged, Z., Hublin, J., Skinner, M., Maxillary molar enamel thickness of Plio-Pleistocene hominins. Journal of Human Evolution. 138(3). Online Article
Milman, E., Daugherty, J., Alemseged, Z., Brennan, K., Lebowicz, L., Visualization of a Juvenile Australopithecus afarensis Specimen: Implications for Functional Foot Anatomy. Journal of Biocommunications. 43(2). Online Article
Du, A., Rowan, J., Wang, S.C., Wood, B. A., Alemseged, Z., Statistical estimates of hominin origination and extinction dates: A case study examining the Australopithecus anamensis-afarensis lineage. Journal of Human Evolution. 138(1). Online Article
2019
Nalley, T.K., Scott, J.E., Ward, C.V., Alemseged, Z., 2019. Comparative morphology and ontogeny of the thoracolumbar transition in great apes, humans, and fossil hominins. Journal of Human Evolution. 134, 102632. Online Article
Habermann, J. M., Alberti, M., Aldeias, V., Alemseged, Z., Archer, W., Bamford, M., … Carvalho, S. (2019). Gorongosa by the sea: First Miocene fossil sites from the Urema Rift, central Mozambique, and their coastal paleoenvironmental and paleoecological contexts. Palaeogeography, Palaeoclimatology, Palaeoecology, 514, 723–738. Online Article
Martinez, F. I., Capelli, C., Ferreira da Silva, M. J., Aldeias, V., Alemseged, Z., Archer, W., … Carvalho, S. (2019). A missing piece of the Papio puzzle: Gorongosa baboon phenostructure and intrageneric relationships. Journal of Human Evolution, 130, 1–20. Online Article
Geraads, D., Barr, W.A., Reed D., Laurin, M., Alemseged, Z., New Remains of Camelus grattardi (Mammalia, Camelidae) from the Plio-Pleistocene of Ethiopia and the Phylogeny of the Genus. Journal of Mammalian Evolution. Online Article
Du, A. & Alemseged, Z., Temporal evidence shows Australopithecus sediba is unlikely to be the ancestor of Homo. Sci. Adv. 5: eaav9038. Online Article
Alemseged, Z., Njau, J., Pobiner, B., and Ndiema, E. Connecting palaeoscientists in eastern Africa and the wider world. Nature Ecology and Evolution, 1. Online Article
Thompson, J.C., Carvalho, S., Marean, C.W., Alemseged, Z. Origins of the human predatory pattern: The transition to large animal exploitation by early hominins. Current Anthropology. 60(1). Online Article
2018
Du, A., Alemseged, Z. Diversity analysis of Plio-Pleistocene large mammal communities in the Omo-Turkana Basin, eastern Africa. J.Hum. Evol. 124: 25-39. Online Article
DeSilva, J.M., Gill, C.M., Prang, T.C., Bredella, M.A., Alemseged, Z. A nearly complete foot from Dikika, Ethiopia and its implications for the ontogeny and function of Australopithecus afarensis. Science Advances, 4(7), eaar7723. Online Article
2017
Ward, C.V., Nalley, T.K., Spoor, F., Tafforeau, P., Alemseged, Z. Thoracic vertebral count and thoracolumbar transition in Australopithecus afarensis. PNAS. 114(23), 6000-6004. Online Article
2016
Wynn, Jonathan G., Reed, K.E., Sponheimer, M., Kimbel, W.H., Alemseged, Z., Bedaso, Z.K., Campisano, C.J. Dietary flexibility of Australopithecus afarensis in the face of paleoecological change during the middle Pliocene: Faunal evidence from Hadar, Ethopia. J. Hum. Evol. 99: 93-106. Online Article
Young, N.M., Capellini, T.D., Roach, N.T., Alemseged, Z. Reply to Almécija: A new direction for reconstructing our last common ancestor with chimpanzees. PNAS 1525673113. Online Article
2015
Young, N.M., Capellini, T.D., Roach, N.T., Alemseged, Z. Reply to Melillo: Woranso-Mille is consistent with an australopithecine shoulder intermediate between African apes and Homo.PNAS 112:E7160. Online Article
Alemseged, Z. Stable isotopes serving as a checkpoint. PNAS 112:12232-12233. Online Article
Reed, D., Barr, W.A., McPherron, S.P., Bobe, R., Geraads, D., Wynn, J.G., Alemseged, Z. Digital data collection in paleoanthropology. Evol. Anth. 24:238-249. Online Article
Young, N.M., Capellini, T.D., Roach, N.T., Alemseged, Z. Fossil hominin shoulders support an African ape-like last common ancestor of humans and chimpanzees. PNAS 112:11829-11834. Online Article
Plummer, T., Ferraro, J.V., Louys, J.C., Hertel, F., Alemseged, Z., Bobe, R., Bishop, L.C. Bovid ecomorphology and hominin paleoenvironments of the Shungura Formation, Lower Omo River Valley, Ethiopia. J. Hum. Evol. 88:108-126. Online Article
Thompson, J.C., McPherron, S.P., Bobe, R., Reed, D., Barr, A., Wynn, J., Marean, C.W., Geraads, D., Alemseged, Z. Taphonomy of fossils from hominin-bearing deposits at Dikika, Ethiopia. J. Hum. Evol. 86:112-135. Online Article
Negash, E., Alemseged, Z., Wynn, J.G., Bedaso, Z.K. Paleodietary reconstruction using stable isotopes and abundance analysis of bovids from the Shungura Formation of South Omo, Ethiopia. J. Hum. Evol. 88:127-136. Online Article
Skinner, M.M., Alemseged, Z., Gaunitz, C., Hublin, J-J. Enamel thickness trends in Plio-Pleistocene hominin mandibular molars. J. Hum. Evol.85:35-45. Online Article
Geraads, D., Alemseged, Z., Bobe, R., Reed, D. Pliocene carnivora (Mammalia) from the Hadar Formation at Dikika, Lower Awash Valley, Ethiopia. J. Afr. Sci. 107:28-35. Online Article
2014
Alemseged, Z. Early Hominins. In: Renfrew, C., Bahn, P. (Ed. 1), The Cambridge World Prehistory. Cambridge University Press, pp. 47-64. Online Article
2013
Sponheimer, M., Alemseged, Z., Cerline, T.E., Grine, F.E., Kimbel, W.H., Leakey, M.G., Lee-Thorp, J.A., Manthi, F.K., Reed, K.E., Wood, B.A., Wynn, J.G. Reply to Fontes-Villalba et al.: On a reluctance to conjecture about animal food consumption. PNAS 110(43):E4056. Online Article
Wynn, J.G., Sponheimer, M., Kimbel, W.H., Alemseged, Z., Reed, K., Bedaso, Z.K., Wilson, J.N. Diet of Australopithecus afarensis from the Pliocene Hadar Formation, Ethiopia. PNAS 110(26):10495-10500. Online Article
Sponheimer, M., Alemseged, Z., Cerling, T.E., Grine, F.G., Kimbel, W.H., Leakey, M.G., Lee-Thorp, J., Manthi, F.K., Reed, K. Wood, B., Wynn, J.G. Isotopic evidence of early hominin diets. PNAS 110(26):10513-10518. Online Article
Alemseged, Z. Australopithecus in Ethiopia. In: Reed, K., Fleagle, J.G., and Leakey, R.E. (Eds.), The Paleobiology of Australopithecus. Springer, Dordrecht, pp. 63-71. Online Article
Smith, T., Alemseged, Z. Reconstructing hominid life history. Nature Ed. Knowl. 4(4):2. Online Article
Bedaso, Z., Wynn, J.G., Alemseged, Z., Geraads, D. Dietary and paleoenvironmental reconstruction using stable isotope analysis of herbivore tooth enamel from the Pliocene locality of Dikika, Ethiopia: implication for Australopithecus afarensis habitat and food resource. J. Hum. Evol.64:21-38. Online Article
2012
Bostelmann, J.E., Bobe, R., Carrasco, G., Alloway, B.V., Santi-Malnis, P., Mancuso, A., Aguero, B., Alemseged, Z., Godoy, Y. The Alto Rio Cisnes fossil fauna (Rio Frias Formation, Early Middle Miocene, Friasian Salma): a keystone and paradigmatic vertebrate assemblage of the South American fossil record. In: Leppe, M., Aravena, J.C., Villa-Martinez, R. (Eds.), Abriendo Ventana al Pasado Libro de Resumenes III Simposio-Paleontologia en Chile. INACH, pp. 42-45. Online Article
Green, D.J., Alemseged, Z. Australopithecus afarensis scapular ontogeny, function, and the role of climbing in human evolution. Science 338:514-517. Online Article
Alemseged, Z., Kiura, P.W., Mbua, E., Njau, J., Pobiner, B. A platform for East African Paleoanthropology: third bi-annual conference of the EAAPP. Evol. Anthropol. 21:89-91. Online Article
2011
McPherron, S.P., Alemseged, Z., Marean, C.W., Wynn, J.G., Reed, D., Geraads, D., Bobe, R., Béarat, H.A. Tool-marked bones from before the Oldowan change the paradigm. PNAS 108:E116. Online Article
Geraads, D., Alemseged, Z., Bobe, R., Reed, D. Enhydriodon dikikae sp. nov. (Carnivora: Mammalia), a gigantic otter from the Pliocene of Dikika, Lower Awash, Ethiopia. J. Vert. Paleontol.31:447-453. Online Article
2010
McPherron, S.P., Alemseged, Z., Marean, C.W., Wynn, J.G., Reed, D., Geraads, D., Bobe, R., Béarat, H. Evidence for stone tool-assisted consumption of animal tissues prior to 3.39 million years ago at Dikika, Ethiopia. Nature 466:857-860. Online Article
Bedaso, Z., Wynn, J.G., Alemseged, Z., Bobe, R., Reed, D. Paleoenvironmental reconstruction of the Asbole fauna (Busidima Formation, Afar, Ethiopia) using stable isotopes. Geobios 43:165-177. Online Article
Geraads, D., Alemseged, Z., Bobe, R., Reed, D. Nyctereutes lockwoodi, n. sp., a new canid (Carnivora: Mammalia) from the Middle Pliocene of Dikika, Lower Awash, Ethiopia. J. Vert. Paleontol.30:981-987. Online Article
2009
Alemseged, Z., Bobe, R. Diet in early hominin species: a paleoenvironmental perspective. In: Hublin, J.J., Richards, M.P. (Eds.), The Evolution of Hominid Diets: Integrating Approaches to the Study of Palaeolithic Subsistence. Springer, Dordrecht, pp. 179-186. Online Article
Njau, J., Mbua, E., Alemseged, Z., Pobiner, B. Second conference of the East African Association for Paleoanthropology and Paleontology: Fifty years after discovery of Zinjanthropus. Evol. Anthropol. 18:235-236. Online Article
2008
Wynn, J. G., Roman, D.C., Alemseged, Z., Reed, D., Geraads, D., Munro, S. Stratigraphy, depositional environments and basin structure of the Hadar and Busidima Formations at Dikika, Ethiopia. In: Quade, J., Wynn, J.G. (Eds.), The Geology of Early Humans in the Horn of Africa: Geological Society of America Special Paper 446, pp. 1-32. Online Article
Mbua, E., Alemseged, Z., Bobe, R. A new association for East African paleoanthropology and paleontology. Evol. Anthropol. 17:123-124. Online Article
2007
Bobe, R., Alemseged, Z., Behrensmeyer, A.K. (Eds.), Hominin Environments in the East African Pliocene: An Assessment of the Faunal Evidence. Springer, Dordrecht. Online Access
Alemseged, Z., Bobe, R., Geraads, D. Comparability of fossil data and its significance for the interpretation of hominin environments: a case study in the lower Omo valley, Ethiopia. In: Bobe, R., Alemseged, Z., Behrensmeyer, A.K. (Eds.), Hominin Environments in the East African Pliocene: An Assessment of the Faunal Evidence. Springer, Dordrecht, pp. 159-181. Online Article
Bobe, R., Behrensmeyer, A.K., Alemseged, Z. Approaches to the analysis of faunal change during the East African Pliocene. In: Bobe, R., Alemseged, Z., Behrensmeyer, A.K. (Eds.), Hominin Environments in the East African Pliocene: An Assessment of the Faunal Evidence. Springer, Dordrecht, pp. 1-24. Online Article
Bobe, R., Behrensmeyer, A.K., Alemseged, Z. Finale and future: investigating faunal evidence for hominin paleoecology in East Africa. In: Bobe, R., Alemseged, Z., Behrensmeyer, A.K. (Eds.), Hominin Environments in the East African Pliocene: An Assessment of the Faunal Evidence. Springer, Dordrecht, pp. 333-345. Online Article
Frost, S.R., Alemseged, Z. Middle Pleistocene fossil Cercopithecidae from Asbole, Afar Region, Ethiopia. J. Hum. Evol. 53:227-259. Online Article
Bobe, R., Alemseged, Z., Behrensmeyer, A.K. (Eds.). Hominin Environments in the East African Pliocene: An Assessment of the Faunal Evidence. Vertebrate Paleobiology and Paleoanthropology Series, Springer, Dordrecht. Online Article
2006
Alemseged, Z., Spoor, F., Kimbel, W.H., Bobe, R., Geraads, D., Reed, D., Wynn, J.G. A juvenile early hominin skeleton from Dikika, Ethiopia. Nature 443:296-301. Online Article
Wynn, J.G., Alemseged, Z., Bobe, R., Geraads, D., Reed, D., Roman, D.C. Geological and palaeontological context of a Pliocene juvenile hominin at Dikika, Ethiopia. Nature 443:332-336. Online Article
2005
Alemseged, Z., Wynn, J.G., Kimbel, W.H., Reed, D., Geraads, D., Bobe, R. A new hominin from the Basal Member of the Hadar Formation, Dikika, Ethiopia and its geological context. J. Hum. Evol. 49:499-514. Online Article
2004
de Lumley, H., Bahain, J.-J. Beyene, Y., Marchal, F., Alemseged, Z. Les formation Plio-Pleistocenes du bassin de l’Omo-Turkana. In: De Lumley, H., Beyene, Y. (Eds.), Les Sites Prehistoriques de la Region de Fejej, Sud-Omo, Ethiopie, dans leurs Contexte Stratigraphique et Paleontologique. Éditions Recherche sur les Civilizations, Paris, pp. 81-107.
Echassoux, A., Moulle, P.E., Desclaux, E., Alemseged, Z. Les faunes Plio-Pleistocene du site de Fejej FJ-1. In: De Lumley, H., Beyene, Y. (Eds.), Les Sites Prehistoriques de la Region de Fejej, Sud-Omo, Ethiopie, dans leurs Contexte Stratigraphique et Paleontologique. Éditions Recherche sur les Civilizations, Paris, pp. 203-340. Online Article
Geraads, D., Alemseged, Z., Reed, D., Wynn, J. The Pleistocene fauna (other than Primates) from Asbole, lower Awash Valley, Ethiopia, and its environmental and biochronological implications. Geobios 37:697-718. Online Article
2003
Alemseged, Z. An integrated approach to taphonomy and faunal change in the Shungura Formation (Ethiopia) and its implication for hominid evolution. J. Hum. Evol. 44:1-28. Online Article
2002
Alemseged, Z., Coppens, Y., Geraads, D. Hominid cranium from Omo: Description and taxonomy of Omo-323-1976-896. Am. J. Phys. Anthropol. 117:103-112. Online Article
Geraads D., Alemseged, Z., Bellon, H. The late Miocene Mammalian fauna of Chorora, Awash basin, Ethiopia: systematics, biochronology and 40K-40Ar ages of associated volcanics. Tertiary Res. 21(1-4):113-122. Online Article
2001
Moullé, P.E., Echassoux, A., Alemseged, Z., Desclaux, E. On the presence of Elephas recki at the Oldowan prehistoric site of Fejej FJ-1 (Ethiopia). International Congress on The World of Elephants, Rome, pp. 122-125. Online Article
2000
Alemseged, Z., Geraads, D. A new Middle Pleistocene Fauna from the Busidima-Telalak region of the Afar, Ethiopia. C. R. Acad. Sci., Paris 331:549-556. Online Article
Alemseged, Z. Les Hominidés plio-pléistocènes de la basse vallée de l’Omo (Éthiopie) et leurs environnements [Plio-pleistocene hominids of the lower Omo valley (Ethiopia) and their environments]. Annales d’Éthiopie, 16(1), 1–9. Online Article
Alemseged, Z. Plio-Pleistocene Hominids and Their Environment in the Lower Omo Valley, Ethiopia. Annls. Ethiopie, 20:1-5.
1999
Alemseged, Z. La basse valle de l’Omo (Ethiopie). In: Berillon, G., Bacon, A.M., Marchal, F., Deloison, Y. (Eds.), Les Australopitheques. Editions Artcom.
1998
Alemseged, Z., Geraads, D. Theropithecus atlanticus (Thomas, 1884) (Primates: Cercopithecidae) from the late Pliocene of Ahl al Oughlam, Casablanca, Morocco. J. Hum. Evol. 34(6):609-621. Online Article
1996
Alemseged, Z., Geraads, D., Coppens, Y., Guillemot, C. Taphonomical and paleoenvironmental study of Omo-33, a late Pliocene hominid locality of the lower Omo basin, Ethiopia. Rev. Paléobiol. 15(2):339-347. Online Article